The associated article can be accessed here, with many more photos.
These photos apparently are a result of film footage taken for a BBC Documentary on Manta Rays: Andrea: Queen of the Mantas
The Andrea in question is biologist Andrea Marshall. Apparently, until recently, it was thought there was a single species of manta ray. She noticed discrepancies in the markings and behaviors of some individuals and was able to describe a new species of manta, Manta alfredi.
